Persian Leopard Bibliography

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Persian Leopard National Conservation and Management Action Plan in Iran Download Endorsement Letter

Conservation Road Map of Cats of Iran Download File

دانلود فایل فارسی نقشه راه حفاظت گربه سانان ایران

Persian Leopard Distribution Studies in Iran

Study Span of 2007-2011: Sanei A., Mousavi M., Kiabi B.H., Masoud M.R., Gord Mardi E., Mohamadi H., Shakiba M., Baran Zehi A., Teimouri M., Raeesi T. 2016. Status assessment of the Persian leopard in Iran. Cat News Special Issue 10: 43-50. Download File

Study Span of 2002-2006: Sanei A. and Zakaria M. 2011. Distribution pattern of the Persian leopard (Panthera pardus saxicolor) in Iran. Asia Life Sciences Supplement 7: 7-18.  Download File

Historical information and baseline knowledge: Sanei A. 2007. Assessment of leopard (Panthera pardus) status in Iran (No.1). Tehran: Sepehr Publication Centre. (in Persian) Download File

The bibliography is in alphabetical order using the author's last name.


Abdollahi S. 2015. Modeling habitat requirements of leopard (Panthera pardus) using genetic algorithm in Golestan National Park. Environmental Resources Research3 (2): 151-162. Download File

Abdoli A., Ghadirian T., Hamidi A.K., Mostafavi H., Moshiri H., Pour’salem S. and Ghoddousi A. 2008. Photo of a Persian leopard from Khaeez area in southern Iran. Cat News 48: 14.   Download File

Askerov E., Talibov T., Manvelyan K., Zazanashvili N., Malkhasyan A., Fatullayev P. and Heidelberg A. 2015. South-Eastern Lesser Caucasus: the most important landscape for conserving the leopard (Panthera pardus) in the Caucasus region (Mammalia: Felidae). Zoology in the Middle East. 61(2): 95-101. Download File

Avgan B., Raza H., Barzani M., and Breitenmoser U. 2016. Do recent leopard Panthera pardus records from northern Iraq and south-eastern Turkey reveal an unknown population nucleus in the region? Zoology in the Middle East 62(2): 95-104. Download File


Babrgir S., Farhadinia M.S. and Moqanaki E.M. 2015. Socio-economic consequences of cattle predation by the Endangered Persian leopard Panthera pardus saxicolor in a Caucasian conflict hotspot, northern Iran. Oryx, 51(1), 124-130. Download File

Bazghandi M. 2016. Economic valuation of endangered wildlife species subjected to trafficking (case study: endangered Persian leopard). Msc thesis. Yazd: Yazd University, School of Natural Resources and Desert studies. (Thesis)

Bernal S.D., Hoby S. and Levionnois O.L. R. 2019. Acute hyperkalaemia in a captive Persian leopard (Panthera pardus saxicolor) immobilised with a ketamine-medetomidine combination. Veterinary Record Case Reports7 (1). Download File


Chalani M. 2005. Analysis of leopard diet in Tandoureh National Park. BSc. thesis. Markazi: Azad University of Arak.   (Thesis)

Cheraghi F., Delavar M.R., Amiraslani F., Alavipanah K., Gurarie E., Jowkar, H., ... and Fagan W.F. 2019. Inter-dependent movements of Asiatic Cheetahs Acinonyx jubatus venaticus and a Persian Leopard Panthera pardus saxicolor in a desert environment in Iran (Mammalia: Felidae). Zoology in the Middle East, 1-10.  Abtsract


Dresser B.L., KramerL., Reece B. and Russell P.T. 1982. Induction of ovulation and successful artificial insemination in a Persian leopard (Panthera pardus saxicolor). Zoo Biol., 1: 55–57. Abstract


Erfanian B., Mirkarimi S.H., Mahini A.S. and Rezaei H.R. 2012. A presence-only habitat suitability model for Persian leopard Panthera pardus saxicolor in Golestan National Park, Iran. Wildlife Biology 19 (2):170-178. Download File


Fadakar D., Rezaei H.R., Hosseini, Sheykhi Ilanloo S., Zamani W. 2012. Detecting Domestic Dog (Canis lupus familiaris) in Diet of Persian Leopard (Panthera pardus saxicolor) Using DNA Tools. The International Journal of Environmental Resources Research 1 (3): 247-252. Download File

Farashi A., and Shariati M. 2018. Evaluation of the role of the national parks for Persian leopard (Panther pardus saxicolor, Pocock 1927) habitat conservation (case study: Tandooreh National Park, Iran). Mammal Research63(4): 425-432.  Download File

Farhadinia M.S., Farahmand H., Gavashelishvili A., Kaboli M., Karami M., Khalili B. and Montazamy S. 2015. Molecular and craniological analysis of leopard, Panthera pardus (Carnivora: Felidae) in Iran: support for a monophyletic clade in Western Asia. Biological Journal of the Linnean Society 114 (4): 721-736.  Download File

Farhadinia M.S., Johnson P.J., Macdonald D.W. and Hunter, L.T. 2018. Anchoring and adjusting amidst humans: Ranging behavior of Persian leopards along the Iran-Turkmenistan borderland. PloS one, 13 (5):e0196602. Download File

Farhadinia M.S., Kaboli M., Karami M. and Farahmand H. 2014. Patterns of sexual dimorphism in the Persian Leopard (Panthera pardus saxicolor) and implications for sex differentiation. Zoology in the Middle East, 60(3): 195-207.  Download File


Gavashelishvili A. and Lukarevskiy V. 2008. Modelling the habitat requirements of leopard Pan­thera pardus in west and central Asia. Journal of Applied Ecology 45: 579-588. Download File

Ghaemi et al. 2012. Study on Helminithic Parasites of Persian leopard in Golestan National Park and Biosphere Reserve of Iran. World Applied Sciences Journal 13 (6): 1471-1473. Download File

Ghoddousi A. 2016. Prey preferences of the Persian leopard and degree of trophic competition with human hunters in Iran. PhD thesis. Germany: Georg-August-Universität Göttingen. (Thesis).

Ghoddousi A., Hamidi Kh. A., Ghadirian T., Ashayeri D. and Khorozyan I. 2010. The status of the endangered Persian leopard Panthera pardus saxicolor in Bamu National Park, Iran. Oryx 4 (4): 551-557. Abstract  

Ghoddousi A., Hamidi Kh. A.H., Ghadirian T., Ashayeri D., Moshiri H. and Khorozyan I. 2008. The status of the Persian leopard in Bamu National Park, Iran. Cat News 49: 10-13.  Download File

Ghoddousi A., Khaleghi Hamidi A., Ghadirian T., Ashayeri D., Hamzehpour M., Moshiri H., Zohrabi H. and Julayi L. 2008b. Territorial marking by Persian leopard (Panthera par­dus saxicolor Pocock, 1927) in Bamu National Park, Iran. Zoology in the Middle East 44: 101-103.


Joslin P. 1990. Leopards in Iran. 1989 International Leopard Studbook: 13-15.  


Kameli A. 2012. Persian leopard habitat suitability assessment using HSI in North Khorasan province. BSc thesis. Zabol: University of Zabol. (Thesis)

Khorozyan, I. 2014. Morphological variation and sexual dimorphism of the common leopard (Panthera pardus) in the Middle East and their implications for species taxonomy and conservation. Mammalian Biology-Zeitschrift für Säugetierkunde 79 (6): 398-405.  Download File

Khorozyan I.G., Malkhasyan A.G., Asmaryan S.G. and Abramov A.V. 2010. Using geographical mapping and occupancy modeling to study the distribution of the critically endangered leopard (Panthera pardus) population in Armenia. In: “Spatial Complexity, Informatics, and Wildlife Conservation” (Cushman S. and Huettmann F., eds.). Tokyo, Springer Publishers: 331-347.  Download File

Khorozyan I. 2010. Leopard Panthera pardus (Linnaeus, 1758). In: Red Data Book of Animals of the Republic of Armenia (Aghasyan A. and Kalashyan M., eds.). Yerevan, Ministry of Nature Protection: 344.

Khorozyan I.G., Malkhasyan A.G. and Abramov A.V. 2008. Presence–absence surveys of prey and their use in predicting leopard (Panthera pardus) densities: a case study from Armenia.Integrative Zoology 3: 322–332.     Download File

Khorozyan I.G. and Abramov A.V. 2007. The leopard, Panthera pardus, (Carnivora: Felidae) and its resilience to human pressure in the Caucasus. Zoology in the Middle East 41: 11-24.  

Khorozyan I.G., Baryshnikov G.F. and Abramov A.V. 2006. Taxonomic status of the leopard, Pan­thera pardus (Carnivora, Felidae) in the Caucasus and adjacent areas. Russian Journal of Theriology 5 (1): 41-52.

Khorozyan I., Malkhasyan A. and Asmaryan S. 2005. The Persian leopard prowls its way to survival. Endangered Species Update 22: 51-60.

Kiabi B.H., Dareshouri B.F., Ghaemi R.A. and Jahanshahi M. 2002. Population status of the Persian leopard (Panthera pardus saxicolor Pocock, 1927) in Iran. Zoology in the Middle East 26: 41-47. Download File


Lukarevsky V., Akkiev M., Askerov E., Agili A., Can E., Gurielidze Z., Kudaktin A.N., Malkhasyan A. and Yarovenko Yu.A. 2007. Status of the leopard in the Caucasus. Cat News Special Issue 2: 15-21.


Mobargha M. 2006. Habitat evaluation of Persian leopard (Panthera pardus saxicolor) in Turan National Park, Iran. MSc thesis. Tehran: Azad University.  (Thesis)

Mowlavi Gh., Marucci G., Mobedi I., Zahabiioon F., Mirjalali H. and Pozio E. 2009.Trichinella britovi in a leopard (Panthera pardus saxicolor) in Iran. Vet Parasitol. 14:164 (2-4) Download File


Namroodi S., Gholami A., Shariat-Bahadori E. 2016. Toxoplasmosis May Lead to Road Kills of Persian Leopards (Panthera pardus saxicolor) in Golestan National Park, Iran. Journal of Wildlife Diseases 52(2):436-438 Abstract


Omidi M., Kaboli M., Karami M., Salman Mahini A. and Hasan Zadeh Kiabi B. 2010. Modelling of the Persian leopard (Panthera pardus saxicolor) habitat suitability in Kolah-Ghazi National Park using ENFA. Science and Environmental Technology 12 (1): 137-148. (in Persian)  Download File


Rozhnov V.V., Lukarevsky V.S. and Sorokin P.A. 2011. Using molecular genetic characteristics under reintroduction of the leopard (Panthera pardus L., 1758) in the Caucasus. Doklady Akademii Nauk 437 (2): 280-285. (in Russian)


Sanei A. 2004. Assessment of leopard status in Iran. BSc. thesis. Tehran: Azad University. (Thesis)

Sanei A. 2018. Modeling regional cumulative effects of land use and land cover variations in relation to persistence of the Persian leopard (Panthera pardus saxicolor) in Iran. PhD Thesis. Malaysia: Universiti Putra Malaysia. (Thesis)

Sanei A. 2007. Assessment of leopard (Panthera pardus) status in Iran (No.1). Tehran: Sepehr Publication Centre. (in Persian) Download File

Sanei A. and Zakaria M. 2008. Distribution of Panthera pardus in Iran in relation to its habitat and climate type. In: 3rd Regional Symposium on Environment and Natural Resources (Saiful A., Norhayati M.O., Shuhaimi A.K. and Zulfahmi A.R., eds.). Selangor, Universiti Kebangsan Malaysia: 54.

Sanei A. and Zakaria M. 2011. Distribution pattern of the Persian leopard (Panthera pardus saxicolor) in Iran. Asia Life Sciences Supplement 7: 7-18.  Download File

Sanei A. and Zakaria M. 2011. Survival of the Persian leopard (Panthera pardus saxicolor) in Iran: primary threats and human-leopard conflicts. Asia Life Sciences Supplement 7: 31-39.  Download File

Sanei A., Zakaria M. and Hermidas Sh. 2011. Prey composition in the Persian leopard distribution range in Iran. Asia Life Sciences Supplement 7: 19-30.  Download File

Sanei A.,  Mousavi  M.,  Mousivand M. and  Zakaria M. 2012. Assessment of the Persian leopard mortality rate in Iran In: Proceedings of UMT 11th International Annual Symposium on Sustainability Science and Management Edited by:B. Arifin, S. N. Afifah Jaafar, S. M. Zain S. Hasan, A. Hassan, H. M. E. Armanto, Ch. Tse Seng, W. Z. Wan Sembok, Sh. Md Sheriff, S. Noordin Mokhtar, F. I. Abdul Wahab, H. Mohd Yusof, A. Mat. Arshad. Universiti Malaysia Terengganu:Malaysia. Download File

Sanei A., Gordmardi E., Jafari B., Absalan H. and Zakaria M. 2013. Persian leopard distribution in relation to human pressures and prey resources in North Khorasan province, Iran In: Proceedings of international forestry graduate students’ conference Edited by: F. S. Lai, R. Halis, S.N. Abu Bakar, S. Ramachandran, C. L. Puan. Faculty of Forestry, Universiti Putra Malaysia: Malaysia.  Download File

Sanei A. and Masoud, M.R. A report to the Persian leopard status and conservation in the Iranian part of the Caucasus eco-region (report no. ALSS/2014/Cau/7-1). Prepared for the Caucasus leopard expert’s workshop, Georgia, 9–10 October 2014. Download File

Sanei A., Mousavi M., Kiabi B.H., Masoud M.R., Gord Mardi E., Mohamadi H., Shakiba M., Baran Zehi A., Teimouri M., Raeesi T. 2016. Status assessment of the Persian leopard in Iran. Cat News Special Issue 10: 43-50. Download File

Sherbafi E. 2010. Analysis of leopard diet in the habitats of Golestan NP. MSc thesis. Tehran: Azad university.  (Thesis)

Sharbafi E., Farhadinia M.S., Rezaie H.R., and Braczkowski A.R. 2016. Prey of the Persian leopard (Panthera pardus saxicolor) in a mixed forest-steppe landscape in northeastern Iran (Mammalia: Felidae). Zoology in the Middle East 62 (1): 1-8. Download File


Taghdisi M., Mohammadi A., Nourani E., Shokri S., Rezaei A. and Kaboli M. 2013. Diet and Habitat Use of the Endangered Persian Leopard (Panthera pardus saxicolor) In Northeastern Iran. Turkish Journal of Zoology 37: 554-561. Download File


Youssefi M.R., Hoseini Sh., Hoseini S.M., Zaheri B.A. and Abouhosseini Tabari A. 2010. First report of Ancylostoma tubaeforme in Persian Leopard (Panthera pardus saxicolor). Iranian Journal of Parasitology 5 (1): 61-63Download File


Zakaria M. and Sanei A. 2011. Conservation and management prospects of the Persian and Malayan leopards.  Asia Life Sciences Supplment 7: 1-5. Download File

منابع فارسی

توکلی و همکاران. 1398. بررسي تفاوت هاي ريخت شناختي جمجمه پلنگ ايراني در زاگرس و نواحي كوهستاني شمال  شمال شرقي (pardus saxicolor Panthera) كشور به روش ريخت سنجي هندسي، محيط زيست طبيعي، منابع طبيعي ايران، ش. 2، ص. 187-198. Download File

رضایی و همکاران. 1395. تعیین پهنای آشیان بوم شناختی غذایی پلنگ ایرانی (Panthera pardus saxicolor) در منطقه حفاظت شده کوه بافق. فصلنامه علمی پژوهشی محیط زیست جانوری. ش. 1، ص. 1-8. Download File

شعاعی و یارمحمدی بربرستانی. 1395. رژيم غذايي پلنگ ايراني (Panthera pardus saxicolor, pocock 1927) در پارک ملي تندوره طي فصول تابستان و پاييز. پژوهش هاي جانوري (زيست شناسي ايران)، ش. 4، ص. 426 - 434.  Download File

صانعی. 1387. بررسی وضعیت پلنگ در ایران (جلد اول). تهران: انتشارات سپهر، 298 ص. Download File

نصیری و همکاران. 1390. تجزیه و تحلیل ژنتیکی و فیلوژنتیکی ناحیه D-Loop از DNA میتوکندری پلنگ ایرانی. مجله بیوتکنولوژی کشاورزی. Download File